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Food habits and consumption rates of common dolphinfish in the eastern Pacific Ocean - Coryphaena hippurus

Fishery Bulletin, April, 2002 by Robert J. Olson, Felipe Galvan-Magana

Table 1
Numbers of common dolphinfish sampled by size, sex, and area with
undigested food in their stomachs (first number) and with empty
stomachs or trace amounts of hard parts (in parentheses).

                               Sex                     Area

Fork length                         Undetermined
(mm)           Males     Females        sex        North   West

417-650       2 (14)     14 (39)     4 (5)         0 (1)    0 (3)
651-800       12 (21)    48 (47)     8 (3)         1 (0)    2 (9)
801-950       24 (29)    51 (70)    11 (7)                 11 (11)
951-1100      26 (25)    15 (21)     3 (1)                  0 (2)
1101-1250      9 (6)     10 (6)      2 (1)         2 (0)
1251-1770      6 (1)      1 (1)      0 (1)
Total         79 (96)   139 (184)   29 (18 (1))    3 (1)   13 (25)

                            Area

Fork length                                         Total no. of
(mm)           East       Southwest     Southeast    dolphinfish

417-650        4 (24)    16 (21)         0 (9)       20 (58)
651-800       14 (11)    37 (47)        14 (4)       68 (71)
801-950       13 (13)    43 (60)        19 (22)      86 (106)
951-1100      10 (10)    19 (19)        15 (16)      44 (47)
1101-1250     10 (5)      5 (1)          4 (7)       21 (13)
1251-1770      1 (1)      2 (1)          4 (1)        7 (3)
Total         52 (64)   122 (150 (1))   56 (59)     246 (299 (1))

(1) Including one dolphinfish with no length or sex data.

Table 2
Taxonomic composition in %W of the prey of common dolphinfish from
five areas of the eastern Pacific Ocean (Fig. 1) and from all areas
combined. Weights do not include hard parts (cephalopod mandibles and
fish otoliths). Prey category codes are as follows: EC = epipelagic
cephalopods, MC = mesopelagic cephalopods, Cr = Crustacea, MiF =
miscellaneous fishes, MsF = mesopelagic fishes, F = flyingfishes,
Co = Coryphaenidae, Ca = Carangidae, G = Gempylidae, A = Auxis spp.,
YT = yellowfin tuna, N = Nomeidae, T = Tetraodontiformes.  [A.sub.i]
is the integral of the function fitted to experimental gastric
evacuation data and is used to calculate consumption rates (Eq. 1).
Area codes are N = north, W = west, E = east, SW = southwest, SE =
southeast.

                                                  Assumed
Taxon                                 Category   [A.sub.i]

Phylum Mollusca                                    4.48
  Class Cephalopoda                                4.48
    Order Teuthoidea                               4.48
      Family Enoploteuthidae            EC         4.48
        Abraliopsis falco               EC         4.48
      Family Mastigoteuthidae           MC         4.48
        Mastigoteuthis spp.             MC         4.48
      Family Ommastrephidae             EC         4.48
        Dosidicus gigas                 EC         4.48
        Sthenoteuthis oualaniensis      EC         4.48
      Family Onychoteuthidae            EC         4.48
        Onychoteuthis banksii           EC         4.48
        Onychoteuthis spp.              EC         4.48
      Family Pholidoteuthidae           EC         4.48
        Pholidoteuthis boschmani        EC         4.48
      Family Thysanoteuthidae           EC         4.48
        Thysanoteuthis rhombus          EC         4.48
    Order Octopoda                      EC         4.48
      Family Tremoctopodidae            EC         4.48
        Tremoctopus violaceus           EC         4.48
      Family Argonautidae               EC         4.48
        Argonauta spp.                  EC         4.48
      Family Bolitaenidae               MC         4.48
        Japetella diaphana              MC         4.48
Phylum Arthropoda                                  3.37
  Class Crustacea                       Cr         3.37
    Order Decapoda                      Cr         3.37
      Family Galatheidae                Cr         3.37
        Pleuroncodes planipes           Cr         3.37
      Family Penaeidae                  Cr         3.37
      Family Portunidae                 Cr         3.37
        Portunus xantusii               Cr         3.37
Phylum Chordata                                    4.12
  Class Osteichthyes                               4.12
    Order Clupeiformes                  MiF        4.12
      Family Engraulidae                MiF        4.12
    Order Stomiiformes                  MsF        2.24
      Family Phosichthyidae             MsF        2.24
        Vinciguerria lucetia            MsF        2.24
    Order Myctophiformes                MsF        2.24
      Family Myctophidae                MsF        2.24
        Benthosema panamense            MsF        2.24
        Myctophum aurolaternatum        MsF        2.24
        Myctophum nitidulum             MsF        2.24
        Symbolophorus spp.              MsF        2.24
    Order Lampridiformes                MsF        5.29
      Family Trachipteridae             MsF        5.29
        Desmodesma polystictum          MsF        5.29
    Order Beloniformes
      Family Belonidae                  MiF        5.29
        Strongylura spp.                MiF        5.29
      Family Exocoetidae                F          4.12
        Cheilopogon furcatus            F          4.12
        Cheilopogon spilonotopterus     F          4.12
        Cheilopogon spp.                F          4.12
        Exocoetus monocirrhus           F          4.12
        Exocoetus volitans              F          4.12
        Exocoetus spp.                  F          4.12
        Hirundichthys speculiger        F          4.12
        Prognichthys spp.               F          4.12
      Family Hemiramphidae              F          4.12
        Oxyporhamphus micropterus       F          4.12
    Order Perciformes
      Family Echeneidae                 MiF        3.77
        Rhombochirus osteochir          MiF        3.77
      Family Coryphaenidae              Co         5.29
        Coryphaena equiselis            Co         5.29
        Coryphaena hippurus             Co         5.29
      Family Carangidae                 Ca         5.29
        Naucrates ductor                Ca         5.29
      Family Gempylidae                 G          5.29
        Gempylus serpens                G          5.29
      Family Scombridae                 MiF        5.29
        Acanthocybium solandri          MiF        5.29
        Auxis spp.                      A          5.29
        Euthynnus lineatus              MiF        5.29
        Thunnus spp.                    YT         5.29
      Family Nomeidae                   N          2.24
        Cubiceps pauciradiatus          N          2.24
    Order Tetraodontiformes             T          3.77
      Family Balistidae                 T          3.77
        Xantichthys mento               T          3.77
      Family Ostraciidae                T          3.77
        Lactoria diaphanum              T          3.77
      Family Tetraodontidae             T          3.77
        Lagocephalus lagocephalus       T          3.77
        Unidentified fishes             MiF        3.77
Total prey weight (g)

                                              Area

Taxon                                   N       W       E

Phylum Mollusca                       16.35    2.54    9.48
  Class Cephalopoda                   16.35    2.54    9.48
    Order Teuthoidea                  16.35    2.54    9.48
      Family Enoploteuthidae                    *
        Abraliopsis falco                       *
      Family Mastigoteuthidae                          0.15
        Mastigoteuthis spp.                            0.15
      Family Ommastrephidae           16.35    2.54    5.47
        Dosidicus gigas                        2.54    0.78
        Sthenoteuthis oualaniensis    16.35            4.69
      Family Onychoteuthidae                           3.26
        Onychoteuthis banksii                          3.26
        Onychoteuthis spp.
      Family Pholidoteuthidae
        Pholidoteuthis boschmani
      Family Thysanoteuthidae                           *
        Thysanoteuthis rhombus                          *
    Order Octopoda                              *       *
      Family Tremoctopodidae
        Tremoctopus violaceus
      Family Argonautidae                       *       *
        Argonauta spp.                          *       *
      Family Bolitaenidae                       *
        Japetella diaphana                      *
Phylum Arthropoda                      1.39            3.26
  Class Crustacea                      1.39            3.26
    Order Decapoda                     1.39            3.26
      Family Galatheidae               1.39
        Pleuroncodes planipes          1.39
      Family Penaeidae                                 0.61
      Family Portunidae                                2.65
        Portunus xantusii                              2.65
Phylum Chordata                       82.26   97.46   87.24
  Class Osteichthyes                  82.26   97.46   87.24
    Order Clupeiformes                                 0.08
      Family Engraulidae                               0.08
    Order Stomiiformes                                 0.48
      Family Phosichthyidae                            0.48
        Vinciguerria lucetia                           0.48
    Order Myctophiformes               0.07    4.26    3.11
      Family Myctophidae               0.07    4.26    3.11
        Benthosema panamense                    *
        Myctophum aurolaternatum                       0.97
        Myctophum nitidulum                             *
        Symbolophorus spp.                     4.26    2.14
    Order Lampridiformes
      Family Trachipteridae
        Desmodesma polystictum
    Order Beloniformes                 1.59   26.50   49.58
      Family Belonidae                                 0.26
        Strongylura spp.                               0.26
      Family Exocoetidae                      16.14   30.77
        Cheilopogon furcatus
        Cheilopogon spilonotopterus
        Cheilopogon spp.                      10.96    0.53
        Exocoetus monocirrhus                         19.19
        Exocoetus volitans                     5.18    5.34
        Exocoetus spp.                                 0.92
        Hirundichthys speculiger                       3.17
        Prognichthys spp.                              1.22
      Family Hemiramphidae             1.59   10.36   18.55
        Oxyporhamphus micropterus      1.59   10.36   18.55
    Order Perciformes                 80.11           32.60
      Family Echeneidae
        Rhombochirus osteochir
      Family Coryphaenidae            80.11            1.43
        Coryphaena equiselis          80.11
        Coryphaena hippurus
      Family Carangidae                                0.34
        Naucrates ductor                               0.34
      Family Gempylidae                               16.53
        Gempylus serpens                              16.53
      Family Scombridae                               13.18
        Acanthocybium solandri                         0.04
        Auxis spp.                                    10.15
        Euthynnus lineatus                             1.50
        Thunnus spp.                                   1.49
      Family Nomeidae                                  1.12
        Cubiceps pauciradiatus                         1.12
    Order Tetraodontiformes            0.49   66.70    1.38
      Family Balistidae                0.21   38.78
        Xantichthys mento                     38.78
      Family Ostraciidae                      25.99
        Lactoria diaphanum                    25.99
      Family Tetraodontidae            0.28    1.93    1.38
        Lagocephalus lagocephalus      0.28    1.93    1.38
        Unidentified fishes
Total prey weight (g)                  1443     985    8459

                                               Area

Taxon                                   SW      SE      All

Phylum Mollusca                        21.69   82.73    32.20
  Class Cephalopoda                    21.69   82.73    32.19
    Order Teuthoidea                   21.66   81.84    31.91
      Family Enoploteuthidae            *        *       *
        Abraliopsis falco               *        *       *
      Family Mastigoteuthidae           0.01             0.04
        Mastigoteuthis spp.             0.01             0.04
      Family Ommastrephidae            21.40   81.84    30.72
        Dosidicus gigas                 5.08   70.73    19.27
        Sthenoteuthis oualaniensis     16.32   11.11    11.45
      Family Onychoteuthidae            0.09             0.91
        Onychoteuthis banksii           0.09             0.91
        Onychoteuthis spp.              *                *
      Family Pholidoteuthidae           *                *
        Pholidoteuthis boschmani        *                *
      Family Thysanoteuthidae           0.17             0.07
        Thysanoteuthis rhombus          0.17             0.07
    Order Octopoda                      0.02    1.16     0.29
      Family Tremoctopodidae                    1.16     0.28
        Tremoctopus violaceus                   1.16     0.28
      Family Argonautidae               0.02     *       0.01
        Argonauta spp.                  0.02     *       0.01
      Family Bolitaenidae               *                *
        Japetella diaphana              *                *
Phylum Arthropoda                       0.01             0.94
  Class Crustacea                       0.01             0.94
    Order Decapoda                      0.01             0.94
      Family Galatheidae                                 0.06
        Pleuroncodes planipes                            0.06
      Family Penaeidae                                   0.16
      Family Portunidae                                  0.71
        Portunus xantusii                                0.71
Phylum Chordata                        78.30   17.00    66.84
  Class Osteichthyes                   78.30   17.00    66.84
    Order Clupeiformes                                   0.02
      Family Engraulidae                                 0.02
    Order Stomiiformes                  4.53             2.01
      Family Phosichthyidae             4.53             2.01
        Vinciguerria lucetia            4.53             2.01
    Order Myctophiformes                *                0.97
      Family Myctophidae                *                0.97
        Benthosema panamense                             *
        Myctophum aurolaternatum                         0.26
        Myctophum nitidulum                              *
        Symbolophorus spp.              *                0.71
    Order Lampridiformes                3.04             1.27
      Family Trachipteridae             3.04             1.27
        Desmodesma polystictum          3.04             1.27
    Order Beloniformes                 31.64    9.29    29.58
      Family Belonidae                          1.34     0.39
        Strongylura spp.                        1.34     0.39
      Family Exocoetidae               28.43    7.95    22.48
        Cheilopogon furcatus            0.66             0.27
        Cheilopogon spilonotopterus     0.12             0.05
        Cheilopogon spp.                3.59    2.94     2.68
        Exocoetus monocirrhus           0.69    1.09     5.69
        Exocoetus volitans             17.64    1.58     9.31
        Exocoetus spp.                  5.04    1.81     2.77
        Hirundichthys speculiger                0.53     0.98
        Prognichthys spp.               0.47             0.52
      Family Hemiramphidae              3.21             6.71
        Oxyporhamphus micropterus       3.21             6.71
    Order Perciformes                  36.32    5.76    28.90
      Family Echeneidae                 2.48             1.03
        Rhombochirus osteochir          2.48             1.03
      Family Coryphaenidae              2.36    2.20     5.08
        Coryphaena equiselis            0.01             3.67
        Coryphaena hippurus             1.23             0.51
      Family Carangidae                 5.48             2.37
        Naucrates ductor                3.88             1.71
      Family Gempylidae                 3.93    0.01     6.07
        Gempylus serpens                3.93    0.01     6.07
      Family Scombridae                15.97    5.54    11.50
        Acanthocybium solandri                           0.01
        Auxis spp.                     15.78             9.29
        Euthynnus lineatus                               0.40
        Thunnus spp.                    0.19    5.54     1.80
      Family Nomeidae                   6.11             2.84
        Cubiceps pauciradiatus          6.11             2.84
    Order Tetraodontiformes             2.77             3.63
      Family Balistidae                                  1.22
        Xantichthys mento                                1.21
      Family Ostraciidae                                 0.81
        Lactoria diaphanum                               0.81
      Family Tetraodontidae             2.77             1.60
        Lagocephalus lagocephalus       2.77             1.60
        Unidentified fishes                     1.95     0.47
Total prey weight (g)                 13,118    7523   31,527

* Only trace quantities of hard parts were present.

Table 3
Taxonomic composition in %N of the prey of common dolphinfish from
five areas of the eastern Pacific Ocean (Fig. 1) and from all areas
combined. Numbers include hard parts (cephalopod mandibles and fish
otoliths) divided by two. Numbers are not complete for some taxa
because sometimes prey could not be enumerated due to being in an
advanced digestion state. Area codes are N = north, W = west,
E = east, SW = southwest, SE = southeast.

                                              Area

Taxon                                   N       W       E

Phylum Mollusca                        2.17   12.36   24.46
  Class Cephalopoda                    2.17   12.36   24.46
    Order Teuthoidea                   2.17    7.87   24.21
      Family Enoploteuthidae                   1.12
        Abraliopsis falco                      1.12
      Family Mastigoteuthidae                          0.24
        Mastigoteuthis spp.                            0.24
      Family Ommastrephidae            2.17    6.74    4.60
        Dosidicus gigas                        6.74    2.91
        Sthenoteuthis oualaniensis     2.17            1.69
      Family Onychoteuthidae                          18.16
        Onychoteuthis banksii                         18.16
        Onychoteuthis spp.
      Family Pholidoteuthidae
        Pholidoteuthis boschmani
      Family Thysanoteuthidae                          0.24
        Thysanoteuthis rhombus                         0.24
    Order Octopoda                             4.49    0.24
      Family Tremoctopodidae
        Tremoctopus violaceus
      Family Argonautidae                      2.25    0.24
        Argonauta spp.                         2.25    0.24
      Family Bolitaenidae                      2.25
        Japetella diaphana                     2.25
Phylum Arthropoda                     69.57           12.38
  Class Crustacea                     69.57           12.38
    Order Decapoda                    69.57           12.38
      Family Galatheidae              69.57
        Pleuroncodes planipes         69.57
      Family Penaeidae                                 1.69
      Family Portunidae                               10.65
        Portunus xantusii                             10.65
Phylum Chordata                        2.17   87.64   62.95
  Class Osteichthyes                   2.17   87.64   62.95
    Order Clupeiformes                                 0.48
      Family Engraulidae                               0.48
    Order Stomiiformes                                  *
      Family Phosichthyidae                             *
        Vinciguerria lucetia                            *
    Order Myctophiformes               2.17   57.30   14.77
      Family Myctophidae               2.17   57.30   14.77
        Benthosema panamense                  16.85
        Myctophum aurolaternatum                       3.39
        Myctophum nitidulum                            3.15
        Symbolophorus spp.                    40.45    8.23
    Order Lampridiformes
      Family Trachipteridae
        Desmodesma polystictum
    Order Beloniformes                 4.35   11.24   36.08
      Family Belonidae                                 0.24
        Strongylura spp.                               0.24
      Family Exocoetidae                       6.74   19.13
        Cheilopogon furcatus
        Cheilopogon spilonotopterus
        Cheilopogon spp.                       2.25    0.48
        Exocoetus monocirrhus                          8.96
        Exocoetus volitans                     4.49    5.33
        Exocoetus spp.                                 1.94
        Hirundichthys speculiger                       1.45
        Prognichthys spp.                              0.97
      Family Hemiramphidae             4.35    4.49   16.71
        Oxyporhamphus micropterus      4.35    4.49   16.71
    Order Perciformes                 17.39           10.41
      Family Echeneidae
        Rhombochirus osteochir
      Family Coryphaenidae            17.39            0.24
        Coryphaena equiselis          17.39
        Coryphaena hippurus
      Family Carangidae                                0.97
        Naucrates ductor                               0.97
      Family Gempylidae                                3.63
        Gempylus serpens                               3.63
      Family Scombridae                                1.69
        Acanthocybium solandri                         0.24
        Auxis spp.                                     0.97
        Euthynnus lineatus                             0.24
        Thunnus spp.                                   0.24
      Family Nomeidae                                  3.87
        Cubiceps pauciradiatus                         3.87
    Order Tetraodontiformes            4.35   19.10    1.21
      Family Balistidae                2.17    2.25
        Xantichthys mento                      2.25
      Family Ostraciidae                      15.73
        Lactoria diaphanum                    15.73
      Family Tetraodontidae            2.17    1.12    1.21
        Lagocephalus lagocephalus      2.17    1.12    1.21
        Unidentified fishes
Total number of prey                     46      89     412

                                              Area

Taxon                                  SW      SE      All

Phylum Mollusca                       19.97   87.35   37.43
  Class Cephalopoda                   19.97   87.35   37.37
    Order Teuthoidea                  16.93   85.25   35.27
      Family Enoploteuthidae           3.62    0.23    1.62
        Abraliopsis falco              3.62    0.23    1.62
      Family Mastigoteuthidae          0.29            0.18
        Mastigoteuthis spp.            0.29            0.18
      Family Ommastrephidae           11.87   85.01   28.20
        Dosidicus gigas                9.99   82.20   26.23
        Sthenoteuthis oualaniensis     1.74    2.81    1.92
      Family Onychoteuthidae           0.72            4.79
        Onychoteuthis banksii          0.58            4.73
        Onychoteuthis spp.             0.14            0.06
      Family Pholidoteuthidae          0.29            0.12
        Pholidoteuthis boschmani       0.29            0.12
      Family Thysanoteuthidae          0.14            0.12
        Thysanoteuthis rhombus         0.14            0.12
    Order Octopoda                     3.04    2.11    2.10
      Family Tremoctopodidae                   0.94    0.24
        Tremoctopus violaceus                  0.94    0.24
      Family Argonautidae              2.75    1.17    1.62
        Argonauta spp.                 2.75    1.17    1.62
      Family Bolitaenidae              0.29            0.24
        Japetella diaphana             0.29            0.24
Phylum Arthropoda                      0.14            5.05
  Class Crustacea                      0.14            5.05
    Order Decapoda                     0.14            5.03
      Family Galatheidae                               1.92
        Pleuroncodes planipes                          1.92
      Family Penaeidae                                 0.42
      Family Portunidae                                2.63
        Portunus xantusii                              2.63
Phylum Chordata                       79.88   12.65   57.31
  Class Osteichthyes                  79.88   12.65   57.31
    Order Clupeiformes                                 0.12
      Family Engraulidae                               0.12
    Order Stomiiformes                41.97           17.37
      Family Phosichthyidae           41.97           17.37
        Vinciguerria lucetia          41.97           17.37
    Order Myctophiformes               0.14            6.83
      Family Myctophidae               0.14            6.83
        Benthosema panamense                           0.90
        Myctophum aurolaternatum                       0.84
        Myctophum nitidulum                            0.78
        Symbolophorus spp.             0.14            4.25
    Order Lampridiformes               0.58            0.24
      Family Trachipteridae            0.58            0.24
        Desmodesma polystictum         0.58            0.24
    Order Beloniformes                29.52   10.77   24.61
      Family Belonidae                         0.23    0.12
        Strongylura spp.                       0.23    0.12
      Family Exocoetidae              26.34   10.54   18.68
        Cheilopogon furcatus           0.29            0.12
        Cheilopogon spilonotopterus    0.29            0.12
        Cheilopogon spp.               1.74    2.34    1.56
        Exocoetus monocirrhus          0.87    0.47    2.69
        Exocoetus volitans            13.31    3.51    7.96
        Exocoetus spp.                 9.55    3.98    5.45
        Hirundichthys speculiger               0.23    0.42
        Prognichthys spp.              0.29            0.36
      Family Hemiramphidae             3.18            5.81
        Oxyporhamphus micropterus      3.18            5.81
    Order Perciformes                  6.80    1.64    6.29
      Family Echeneidae                0.29            0.12
        Rhombochirus osteochir         0.29            0.12
      Family Coryphaenidae             0.72    0.47    0.96
        Coryphaena equiselis           0.14            0.54
        Coryphaena hippurus            0.29            0.12
      Family Carangidae                1.88            1.02
        Naucrates ductor               1.45            0.84
      Family Gempylidae                1.01    0.47    1.44
        Gempylus serpens               1.01    0.47    1.44
      Family Scombridae                1.88    0.70    1.38
        Acanthocybium solandri                         0.06
        Auxis spp.                     1.74            0.96
        Euthynnus lineatus                             0.06
        Thunnus spp.                   0.14    0.70    0.30
      Family Nomeidae                  1.01            1.38
        Cubiceps pauciradiatus         1.01            1.38
    Order Tetraodontiformes            0.72            1.74
      Family Balistidae                                0.18
        Xantichthys mento                              0.12
      Family Ostraciidae                               0.84
        Lactoria diaphanum                             0.84
      Family Tetraodontidae            0.72            0.72
        Lagocephalus lagocephalus      0.72            0.72
        Unidentified fishes            0.14    0.23    0.12
Total number of prey                    691     427    1665

* Individual prey could not be enumerated.

Table 4
Taxonomic composition in %O of the prey of common dolphinfish from
five areas of the eastern Pacific Ocean (Fig. 1) and from all areas
combined. The occurrence of hard parts (cephalopod mandibles and
fish otoliths) was included. Area codes are N = north, W = west,
E = east, SW = southwest, SE = southeast.

                                             Area

Taxon                                  N       W       E

Phylum Mollusca                       25.0   15.79   15.52
  Class Cephalopoda                   25.0   15.79   15.52
    Order Teuthoidea                  25.0   10.53   15.52
      Family Enoploteuthidae                  2.63
        Abraliopsis falco                     2.63
      Family Mastigoteuthidae                         0.86
        Mastigoteuthis spp.                           0.86
      Family Ommastrephidae           25.0    7.89    8.62
        Dosidicus gigas                       7.89    6.03
        Sthenoteuthis oualaniensis    25.0            4.31
      Family Onychoteuthidae                          9.48
        Onychoteuthis banksii                         9.48
        Onychoteuthis spp.
      Family Pholidoteuthidae
        Pholidoteuthis boschmani
      Family Thysanoteuthidae                         0.86
        Thysanoteuthis rhombus                        0.86
    Order Octopoda                            7.89    0.86
      Family Tremoctopodidae
        Tremoctopus violaceus
      Family Argonautidae                     5.26    0.86
        Argonauta spp.                        5.26    0.86
      Family Bolitaenidae                     2.63
        Japetella diaphana                    2.63
Phylum Arthropoda                     25.0            8.62
  Class Crustacea                     25.0            8.62
    Order Decapoda                    25.0            7.76
      Family Galatheidae              25.0
        Pleuroncodes planipes         25.0
      Family Penaeidae                                4.31
      Family Portunidae                               3.45
        Portunus xantusii                             3.45
Phylum Chordata                       50.0   36.84   43.10
  Class Osteichthyes                  50.0   36.84   43.10
    Order Clupeiformes                                0.86
      Family Engraulidae                              0.86
    Order Stomiiformes                                0.86
      Family Phosichthyidae                           0.86
        Vinciguerria lucetia                          0.86
    Order Myctophiformes              25.0   10.53    6.90
      Family Myctophidae              25.0   10.53    6.90
        Benthosema panamense                  2.63
        Myctophum aurolaternatum                      6.03
        Myctophum nitidulum                           0.86
        Symbolophorus spp.                    7.89    3.45
    Order Lampridiformes
      Family Trachipteridae
        Desmodesma polystictum
    Order Beloniformes                25.0   13.16   36.21
      Family Belonidae                                0.86
        Strongylura spp.                              0.86
      Family Exocoetidae                     13.16   25.86
        Cheilopogon furcatus
        Cheilopogon spilonotopterus
        Cheilopogon spp.                      5.26    1.72
        Exocoetus monocirrhus                        12.07
        Exocoetus volitans                    7.89    6.90
        Exocoetus spp.                                2.59
        Hirundichthys speculiger                      1.72
        Prognichthys spp.                             1.72
      Family Hemiramphidae            25.0    2.63   24.14
        Oxyporhamphus micropterus     25.0    2.63   24.14
    Order Perciformes                 50.0           17.24
      Family Echeneidae
        Rhombochirus osteochir
      Family Coryphaenidae            50.0            0.86
        Coryphaena equiselis          50.0
        Coryphaena hippurus
      Family Carangidae                               1.72
        Naucrates ductor                              1.72
      Family Gempylidae                               7.76
        Gempylus serpens                              7.76
      Family Scombridae                               5.17
        Acanthocybium solandri                        0.86
        Auxis spp.                                    2.59
        Euthynnus lineatus                            0.86
        Thunnus spp.                                  0.86
      Family Nomeidae                                 6.90
        Cubiceps pauciradiatus                        6.90
    Order Tetraodontiformes           25.0   18.42    3.45
      Family Balistidae               25.0    5.26
        Xantichthys mento                     5.26
      Family Ostraciidae                     10.53
        Lactoria diaphanum                   10.53
      Family Tetraodontidae           25.0    2.63    3.45
        Lagocephalus lagocephalus     25.0    2.63    3.45
        Unidentified fishes
Total number of samples                  4      38     116

                                              Area

Taxon                                  SW      SE      All

Phylum Mollusca                       14.34   45.22   21.28
  Class Cephalopoda                   13.97   45.22   21.10
    Order Teuthoidea                  12.13   43.48   19.45
      Family Enoploteuthidae           0.37    0.87    0.55
        Abraliopsis falco              0.37    0.87    0.55
      Family Mastigoteuthidae          0.74            0.55
        Mastigoteuthis spp.            0.74            0.55
      Family Ommastrephidae           10.29   43.48   16.88
        Dosidicus gigas                7.35   37.39   13.39
        Sthenoteuthis oualaniensis     2.57    8.70    4.22
      Family Onychoteuthidae           1.84            2.94
        Onychoteuthis banksii          1.47            2.75
        Onychoteuthis spp.             0.37            0.18
      Family Pholidoteuthidae          0.74            0.37
        Pholidoteuthis boschmani       0.74            0.37
      Family Thysanoteuthidae          0.37            0.37
        Thysanoteuthis rhombus         0.37            0.37
    Order Octopoda                     3.68    5.22    3.67
      Family Tremoctopodidae                   3.48    0.73
        Tremoctopus violaceus                  3.48    0.73
      Family Argonautidae              3.68    1.74    2.75
        Argonauta spp.                 3.68    1.74    2.75
      Family Bolitaenidae              0.74            0.55
        Japetella diaphana             0.74            0.55
Phylum Arthropoda                      1.10            2.75
  Class Crustacea                      1.10            2.75
    Order Decapoda                     0.37            2.02
      Family Galatheidae                               0.18
        Pleuroncodes planipes                          0.18
      Family Penaeidae                                 0.92
      Family Portunidae                                0.73
        Portunus xantusii                              0.73
Phylum Chordata                       42.28   25.22   38.53
  Class Osteichthyes                  42.28   25.22   38.53
    Order Clupeiformes                                 0.18
      Family Engraulidae                               0.18
    Order Stomiiformes                 2.21            1.28
      Family Phosichthyidae            2.21            1.28
        Vinciguerria lucetia           2.21            1.28
    Order Myctophiformes               0.37            2.57
      Family Myctophidae               0.37            2.57
        Benthosema panamense                           0.18
        Myctophum aurolaternatum                       1.28
        Myctophum nitidulum                            0.18
        Symbolophorus spp.             0.37            1.47
    Order Lampridiformes               0.74            0.37
      Family Trachipteridae            0.74            0.37
        Desmodesma polystictum         0.74            0.37
    Order Beloniformes                32.35   20.00   29.17
      Family Belonidae                         0.87    0.37
        Strongylura spp.                       0.87    0.37
      Family Exocoetidae              28.68   19.13   24.77
        Cheilopogon furcatus           0.74            0.37
        Cheilopogon spilonotopterus    0.37            0.18
        Cheilopogon spp.               4.04    4.35    3.67
        Exocoetus monocirrhus          0.74    0.87    3.12
        Exocoetus volitans            18.01    6.09   12.29
        Exocoetus spp.                 6.99    9.57    6.06
        Hirundichthys speculiger               0.87    0.55
        Prognichthys spp.              0.74            0.73
      Family Hemiramphidae             6.99            8.99
        Oxyporhamphus micropterus      6.99            8.99
    Order Perciformes                 11.40    5.22   10.83
      Family Echeneidae                0.37            0.18
        Rhombochirus osteochir         0.37            0.18
      Family Coryphaenidae             1.84    1.74    1.83
        Coryphaena equiselis           0.37            0.55
        Coryphaena hippurus            0.74            0.37
      Family Carangidae                1.84            1.28
        Naucrates ductor               1.47            1.10
      Family Gempylidae                1.47    0.87    2.57
        Gempylus serpens               1.47    0.87    2.57
      Family Scombridae                4.41    2.61    3.85
        Acanthocybium solandri                         0.18
        Auxis spp.                     4.04            2.57
        Euthynnus lineatus                             0.18
        Thunnus spp.                   0.37    2.61    0.92
      Family Nomeidae                  1.84            2.39
        Cubiceps pauciradiatus         1.84            2.39
    Order Tetraodontiformes            1.47            2.94
      Family Balistidae                                0.55
        Xantichthys mento                              0.37
      Family Ostraciidae                               0.73
        Lactoria diaphanum                             0.73
      Family Tetraodontidae            1.47            1.83
        Lagocephalus lagocephalus      1.47            1.83
        Unidentified fishes            0.74    0.87    1.10
Total number of samples                 272     115     545
 

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