Food habits and consumption rates of common dolphinfish in the eastern Pacific Ocean - Coryphaena hippurus
Table 1
Numbers of common dolphinfish sampled by size, sex, and area with
undigested food in their stomachs (first number) and with empty
stomachs or trace amounts of hard parts (in parentheses).
Sex Area
Fork length Undetermined
(mm) Males Females sex North West
417-650 2 (14) 14 (39) 4 (5) 0 (1) 0 (3)
651-800 12 (21) 48 (47) 8 (3) 1 (0) 2 (9)
801-950 24 (29) 51 (70) 11 (7) 11 (11)
951-1100 26 (25) 15 (21) 3 (1) 0 (2)
1101-1250 9 (6) 10 (6) 2 (1) 2 (0)
1251-1770 6 (1) 1 (1) 0 (1)
Total 79 (96) 139 (184) 29 (18 (1)) 3 (1) 13 (25)
Area
Fork length Total no. of
(mm) East Southwest Southeast dolphinfish
417-650 4 (24) 16 (21) 0 (9) 20 (58)
651-800 14 (11) 37 (47) 14 (4) 68 (71)
801-950 13 (13) 43 (60) 19 (22) 86 (106)
951-1100 10 (10) 19 (19) 15 (16) 44 (47)
1101-1250 10 (5) 5 (1) 4 (7) 21 (13)
1251-1770 1 (1) 2 (1) 4 (1) 7 (3)
Total 52 (64) 122 (150 (1)) 56 (59) 246 (299 (1))
(1) Including one dolphinfish with no length or sex data.
Table 2
Taxonomic composition in %W of the prey of common dolphinfish from
five areas of the eastern Pacific Ocean (Fig. 1) and from all areas
combined. Weights do not include hard parts (cephalopod mandibles and
fish otoliths). Prey category codes are as follows: EC = epipelagic
cephalopods, MC = mesopelagic cephalopods, Cr = Crustacea, MiF =
miscellaneous fishes, MsF = mesopelagic fishes, F = flyingfishes,
Co = Coryphaenidae, Ca = Carangidae, G = Gempylidae, A = Auxis spp.,
YT = yellowfin tuna, N = Nomeidae, T = Tetraodontiformes. [A.sub.i]
is the integral of the function fitted to experimental gastric
evacuation data and is used to calculate consumption rates (Eq. 1).
Area codes are N = north, W = west, E = east, SW = southwest, SE =
southeast.
Assumed
Taxon Category [A.sub.i]
Phylum Mollusca 4.48
Class Cephalopoda 4.48
Order Teuthoidea 4.48
Family Enoploteuthidae EC 4.48
Abraliopsis falco EC 4.48
Family Mastigoteuthidae MC 4.48
Mastigoteuthis spp. MC 4.48
Family Ommastrephidae EC 4.48
Dosidicus gigas EC 4.48
Sthenoteuthis oualaniensis EC 4.48
Family Onychoteuthidae EC 4.48
Onychoteuthis banksii EC 4.48
Onychoteuthis spp. EC 4.48
Family Pholidoteuthidae EC 4.48
Pholidoteuthis boschmani EC 4.48
Family Thysanoteuthidae EC 4.48
Thysanoteuthis rhombus EC 4.48
Order Octopoda EC 4.48
Family Tremoctopodidae EC 4.48
Tremoctopus violaceus EC 4.48
Family Argonautidae EC 4.48
Argonauta spp. EC 4.48
Family Bolitaenidae MC 4.48
Japetella diaphana MC 4.48
Phylum Arthropoda 3.37
Class Crustacea Cr 3.37
Order Decapoda Cr 3.37
Family Galatheidae Cr 3.37
Pleuroncodes planipes Cr 3.37
Family Penaeidae Cr 3.37
Family Portunidae Cr 3.37
Portunus xantusii Cr 3.37
Phylum Chordata 4.12
Class Osteichthyes 4.12
Order Clupeiformes MiF 4.12
Family Engraulidae MiF 4.12
Order Stomiiformes MsF 2.24
Family Phosichthyidae MsF 2.24
Vinciguerria lucetia MsF 2.24
Order Myctophiformes MsF 2.24
Family Myctophidae MsF 2.24
Benthosema panamense MsF 2.24
Myctophum aurolaternatum MsF 2.24
Myctophum nitidulum MsF 2.24
Symbolophorus spp. MsF 2.24
Order Lampridiformes MsF 5.29
Family Trachipteridae MsF 5.29
Desmodesma polystictum MsF 5.29
Order Beloniformes
Family Belonidae MiF 5.29
Strongylura spp. MiF 5.29
Family Exocoetidae F 4.12
Cheilopogon furcatus F 4.12
Cheilopogon spilonotopterus F 4.12
Cheilopogon spp. F 4.12
Exocoetus monocirrhus F 4.12
Exocoetus volitans F 4.12
Exocoetus spp. F 4.12
Hirundichthys speculiger F 4.12
Prognichthys spp. F 4.12
Family Hemiramphidae F 4.12
Oxyporhamphus micropterus F 4.12
Order Perciformes
Family Echeneidae MiF 3.77
Rhombochirus osteochir MiF 3.77
Family Coryphaenidae Co 5.29
Coryphaena equiselis Co 5.29
Coryphaena hippurus Co 5.29
Family Carangidae Ca 5.29
Naucrates ductor Ca 5.29
Family Gempylidae G 5.29
Gempylus serpens G 5.29
Family Scombridae MiF 5.29
Acanthocybium solandri MiF 5.29
Auxis spp. A 5.29
Euthynnus lineatus MiF 5.29
Thunnus spp. YT 5.29
Family Nomeidae N 2.24
Cubiceps pauciradiatus N 2.24
Order Tetraodontiformes T 3.77
Family Balistidae T 3.77
Xantichthys mento T 3.77
Family Ostraciidae T 3.77
Lactoria diaphanum T 3.77
Family Tetraodontidae T 3.77
Lagocephalus lagocephalus T 3.77
Unidentified fishes MiF 3.77
Total prey weight (g)
Area
Taxon N W E
Phylum Mollusca 16.35 2.54 9.48
Class Cephalopoda 16.35 2.54 9.48
Order Teuthoidea 16.35 2.54 9.48
Family Enoploteuthidae *
Abraliopsis falco *
Family Mastigoteuthidae 0.15
Mastigoteuthis spp. 0.15
Family Ommastrephidae 16.35 2.54 5.47
Dosidicus gigas 2.54 0.78
Sthenoteuthis oualaniensis 16.35 4.69
Family Onychoteuthidae 3.26
Onychoteuthis banksii 3.26
Onychoteuthis spp.
Family Pholidoteuthidae
Pholidoteuthis boschmani
Family Thysanoteuthidae *
Thysanoteuthis rhombus *
Order Octopoda * *
Family Tremoctopodidae
Tremoctopus violaceus
Family Argonautidae * *
Argonauta spp. * *
Family Bolitaenidae *
Japetella diaphana *
Phylum Arthropoda 1.39 3.26
Class Crustacea 1.39 3.26
Order Decapoda 1.39 3.26
Family Galatheidae 1.39
Pleuroncodes planipes 1.39
Family Penaeidae 0.61
Family Portunidae 2.65
Portunus xantusii 2.65
Phylum Chordata 82.26 97.46 87.24
Class Osteichthyes 82.26 97.46 87.24
Order Clupeiformes 0.08
Family Engraulidae 0.08
Order Stomiiformes 0.48
Family Phosichthyidae 0.48
Vinciguerria lucetia 0.48
Order Myctophiformes 0.07 4.26 3.11
Family Myctophidae 0.07 4.26 3.11
Benthosema panamense *
Myctophum aurolaternatum 0.97
Myctophum nitidulum *
Symbolophorus spp. 4.26 2.14
Order Lampridiformes
Family Trachipteridae
Desmodesma polystictum
Order Beloniformes 1.59 26.50 49.58
Family Belonidae 0.26
Strongylura spp. 0.26
Family Exocoetidae 16.14 30.77
Cheilopogon furcatus
Cheilopogon spilonotopterus
Cheilopogon spp. 10.96 0.53
Exocoetus monocirrhus 19.19
Exocoetus volitans 5.18 5.34
Exocoetus spp. 0.92
Hirundichthys speculiger 3.17
Prognichthys spp. 1.22
Family Hemiramphidae 1.59 10.36 18.55
Oxyporhamphus micropterus 1.59 10.36 18.55
Order Perciformes 80.11 32.60
Family Echeneidae
Rhombochirus osteochir
Family Coryphaenidae 80.11 1.43
Coryphaena equiselis 80.11
Coryphaena hippurus
Family Carangidae 0.34
Naucrates ductor 0.34
Family Gempylidae 16.53
Gempylus serpens 16.53
Family Scombridae 13.18
Acanthocybium solandri 0.04
Auxis spp. 10.15
Euthynnus lineatus 1.50
Thunnus spp. 1.49
Family Nomeidae 1.12
Cubiceps pauciradiatus 1.12
Order Tetraodontiformes 0.49 66.70 1.38
Family Balistidae 0.21 38.78
Xantichthys mento 38.78
Family Ostraciidae 25.99
Lactoria diaphanum 25.99
Family Tetraodontidae 0.28 1.93 1.38
Lagocephalus lagocephalus 0.28 1.93 1.38
Unidentified fishes
Total prey weight (g) 1443 985 8459
Area
Taxon SW SE All
Phylum Mollusca 21.69 82.73 32.20
Class Cephalopoda 21.69 82.73 32.19
Order Teuthoidea 21.66 81.84 31.91
Family Enoploteuthidae * * *
Abraliopsis falco * * *
Family Mastigoteuthidae 0.01 0.04
Mastigoteuthis spp. 0.01 0.04
Family Ommastrephidae 21.40 81.84 30.72
Dosidicus gigas 5.08 70.73 19.27
Sthenoteuthis oualaniensis 16.32 11.11 11.45
Family Onychoteuthidae 0.09 0.91
Onychoteuthis banksii 0.09 0.91
Onychoteuthis spp. * *
Family Pholidoteuthidae * *
Pholidoteuthis boschmani * *
Family Thysanoteuthidae 0.17 0.07
Thysanoteuthis rhombus 0.17 0.07
Order Octopoda 0.02 1.16 0.29
Family Tremoctopodidae 1.16 0.28
Tremoctopus violaceus 1.16 0.28
Family Argonautidae 0.02 * 0.01
Argonauta spp. 0.02 * 0.01
Family Bolitaenidae * *
Japetella diaphana * *
Phylum Arthropoda 0.01 0.94
Class Crustacea 0.01 0.94
Order Decapoda 0.01 0.94
Family Galatheidae 0.06
Pleuroncodes planipes 0.06
Family Penaeidae 0.16
Family Portunidae 0.71
Portunus xantusii 0.71
Phylum Chordata 78.30 17.00 66.84
Class Osteichthyes 78.30 17.00 66.84
Order Clupeiformes 0.02
Family Engraulidae 0.02
Order Stomiiformes 4.53 2.01
Family Phosichthyidae 4.53 2.01
Vinciguerria lucetia 4.53 2.01
Order Myctophiformes * 0.97
Family Myctophidae * 0.97
Benthosema panamense *
Myctophum aurolaternatum 0.26
Myctophum nitidulum *
Symbolophorus spp. * 0.71
Order Lampridiformes 3.04 1.27
Family Trachipteridae 3.04 1.27
Desmodesma polystictum 3.04 1.27
Order Beloniformes 31.64 9.29 29.58
Family Belonidae 1.34 0.39
Strongylura spp. 1.34 0.39
Family Exocoetidae 28.43 7.95 22.48
Cheilopogon furcatus 0.66 0.27
Cheilopogon spilonotopterus 0.12 0.05
Cheilopogon spp. 3.59 2.94 2.68
Exocoetus monocirrhus 0.69 1.09 5.69
Exocoetus volitans 17.64 1.58 9.31
Exocoetus spp. 5.04 1.81 2.77
Hirundichthys speculiger 0.53 0.98
Prognichthys spp. 0.47 0.52
Family Hemiramphidae 3.21 6.71
Oxyporhamphus micropterus 3.21 6.71
Order Perciformes 36.32 5.76 28.90
Family Echeneidae 2.48 1.03
Rhombochirus osteochir 2.48 1.03
Family Coryphaenidae 2.36 2.20 5.08
Coryphaena equiselis 0.01 3.67
Coryphaena hippurus 1.23 0.51
Family Carangidae 5.48 2.37
Naucrates ductor 3.88 1.71
Family Gempylidae 3.93 0.01 6.07
Gempylus serpens 3.93 0.01 6.07
Family Scombridae 15.97 5.54 11.50
Acanthocybium solandri 0.01
Auxis spp. 15.78 9.29
Euthynnus lineatus 0.40
Thunnus spp. 0.19 5.54 1.80
Family Nomeidae 6.11 2.84
Cubiceps pauciradiatus 6.11 2.84
Order Tetraodontiformes 2.77 3.63
Family Balistidae 1.22
Xantichthys mento 1.21
Family Ostraciidae 0.81
Lactoria diaphanum 0.81
Family Tetraodontidae 2.77 1.60
Lagocephalus lagocephalus 2.77 1.60
Unidentified fishes 1.95 0.47
Total prey weight (g) 13,118 7523 31,527
* Only trace quantities of hard parts were present.
Table 3
Taxonomic composition in %N of the prey of common dolphinfish from
five areas of the eastern Pacific Ocean (Fig. 1) and from all areas
combined. Numbers include hard parts (cephalopod mandibles and fish
otoliths) divided by two. Numbers are not complete for some taxa
because sometimes prey could not be enumerated due to being in an
advanced digestion state. Area codes are N = north, W = west,
E = east, SW = southwest, SE = southeast.
Area
Taxon N W E
Phylum Mollusca 2.17 12.36 24.46
Class Cephalopoda 2.17 12.36 24.46
Order Teuthoidea 2.17 7.87 24.21
Family Enoploteuthidae 1.12
Abraliopsis falco 1.12
Family Mastigoteuthidae 0.24
Mastigoteuthis spp. 0.24
Family Ommastrephidae 2.17 6.74 4.60
Dosidicus gigas 6.74 2.91
Sthenoteuthis oualaniensis 2.17 1.69
Family Onychoteuthidae 18.16
Onychoteuthis banksii 18.16
Onychoteuthis spp.
Family Pholidoteuthidae
Pholidoteuthis boschmani
Family Thysanoteuthidae 0.24
Thysanoteuthis rhombus 0.24
Order Octopoda 4.49 0.24
Family Tremoctopodidae
Tremoctopus violaceus
Family Argonautidae 2.25 0.24
Argonauta spp. 2.25 0.24
Family Bolitaenidae 2.25
Japetella diaphana 2.25
Phylum Arthropoda 69.57 12.38
Class Crustacea 69.57 12.38
Order Decapoda 69.57 12.38
Family Galatheidae 69.57
Pleuroncodes planipes 69.57
Family Penaeidae 1.69
Family Portunidae 10.65
Portunus xantusii 10.65
Phylum Chordata 2.17 87.64 62.95
Class Osteichthyes 2.17 87.64 62.95
Order Clupeiformes 0.48
Family Engraulidae 0.48
Order Stomiiformes *
Family Phosichthyidae *
Vinciguerria lucetia *
Order Myctophiformes 2.17 57.30 14.77
Family Myctophidae 2.17 57.30 14.77
Benthosema panamense 16.85
Myctophum aurolaternatum 3.39
Myctophum nitidulum 3.15
Symbolophorus spp. 40.45 8.23
Order Lampridiformes
Family Trachipteridae
Desmodesma polystictum
Order Beloniformes 4.35 11.24 36.08
Family Belonidae 0.24
Strongylura spp. 0.24
Family Exocoetidae 6.74 19.13
Cheilopogon furcatus
Cheilopogon spilonotopterus
Cheilopogon spp. 2.25 0.48
Exocoetus monocirrhus 8.96
Exocoetus volitans 4.49 5.33
Exocoetus spp. 1.94
Hirundichthys speculiger 1.45
Prognichthys spp. 0.97
Family Hemiramphidae 4.35 4.49 16.71
Oxyporhamphus micropterus 4.35 4.49 16.71
Order Perciformes 17.39 10.41
Family Echeneidae
Rhombochirus osteochir
Family Coryphaenidae 17.39 0.24
Coryphaena equiselis 17.39
Coryphaena hippurus
Family Carangidae 0.97
Naucrates ductor 0.97
Family Gempylidae 3.63
Gempylus serpens 3.63
Family Scombridae 1.69
Acanthocybium solandri 0.24
Auxis spp. 0.97
Euthynnus lineatus 0.24
Thunnus spp. 0.24
Family Nomeidae 3.87
Cubiceps pauciradiatus 3.87
Order Tetraodontiformes 4.35 19.10 1.21
Family Balistidae 2.17 2.25
Xantichthys mento 2.25
Family Ostraciidae 15.73
Lactoria diaphanum 15.73
Family Tetraodontidae 2.17 1.12 1.21
Lagocephalus lagocephalus 2.17 1.12 1.21
Unidentified fishes
Total number of prey 46 89 412
Area
Taxon SW SE All
Phylum Mollusca 19.97 87.35 37.43
Class Cephalopoda 19.97 87.35 37.37
Order Teuthoidea 16.93 85.25 35.27
Family Enoploteuthidae 3.62 0.23 1.62
Abraliopsis falco 3.62 0.23 1.62
Family Mastigoteuthidae 0.29 0.18
Mastigoteuthis spp. 0.29 0.18
Family Ommastrephidae 11.87 85.01 28.20
Dosidicus gigas 9.99 82.20 26.23
Sthenoteuthis oualaniensis 1.74 2.81 1.92
Family Onychoteuthidae 0.72 4.79
Onychoteuthis banksii 0.58 4.73
Onychoteuthis spp. 0.14 0.06
Family Pholidoteuthidae 0.29 0.12
Pholidoteuthis boschmani 0.29 0.12
Family Thysanoteuthidae 0.14 0.12
Thysanoteuthis rhombus 0.14 0.12
Order Octopoda 3.04 2.11 2.10
Family Tremoctopodidae 0.94 0.24
Tremoctopus violaceus 0.94 0.24
Family Argonautidae 2.75 1.17 1.62
Argonauta spp. 2.75 1.17 1.62
Family Bolitaenidae 0.29 0.24
Japetella diaphana 0.29 0.24
Phylum Arthropoda 0.14 5.05
Class Crustacea 0.14 5.05
Order Decapoda 0.14 5.03
Family Galatheidae 1.92
Pleuroncodes planipes 1.92
Family Penaeidae 0.42
Family Portunidae 2.63
Portunus xantusii 2.63
Phylum Chordata 79.88 12.65 57.31
Class Osteichthyes 79.88 12.65 57.31
Order Clupeiformes 0.12
Family Engraulidae 0.12
Order Stomiiformes 41.97 17.37
Family Phosichthyidae 41.97 17.37
Vinciguerria lucetia 41.97 17.37
Order Myctophiformes 0.14 6.83
Family Myctophidae 0.14 6.83
Benthosema panamense 0.90
Myctophum aurolaternatum 0.84
Myctophum nitidulum 0.78
Symbolophorus spp. 0.14 4.25
Order Lampridiformes 0.58 0.24
Family Trachipteridae 0.58 0.24
Desmodesma polystictum 0.58 0.24
Order Beloniformes 29.52 10.77 24.61
Family Belonidae 0.23 0.12
Strongylura spp. 0.23 0.12
Family Exocoetidae 26.34 10.54 18.68
Cheilopogon furcatus 0.29 0.12
Cheilopogon spilonotopterus 0.29 0.12
Cheilopogon spp. 1.74 2.34 1.56
Exocoetus monocirrhus 0.87 0.47 2.69
Exocoetus volitans 13.31 3.51 7.96
Exocoetus spp. 9.55 3.98 5.45
Hirundichthys speculiger 0.23 0.42
Prognichthys spp. 0.29 0.36
Family Hemiramphidae 3.18 5.81
Oxyporhamphus micropterus 3.18 5.81
Order Perciformes 6.80 1.64 6.29
Family Echeneidae 0.29 0.12
Rhombochirus osteochir 0.29 0.12
Family Coryphaenidae 0.72 0.47 0.96
Coryphaena equiselis 0.14 0.54
Coryphaena hippurus 0.29 0.12
Family Carangidae 1.88 1.02
Naucrates ductor 1.45 0.84
Family Gempylidae 1.01 0.47 1.44
Gempylus serpens 1.01 0.47 1.44
Family Scombridae 1.88 0.70 1.38
Acanthocybium solandri 0.06
Auxis spp. 1.74 0.96
Euthynnus lineatus 0.06
Thunnus spp. 0.14 0.70 0.30
Family Nomeidae 1.01 1.38
Cubiceps pauciradiatus 1.01 1.38
Order Tetraodontiformes 0.72 1.74
Family Balistidae 0.18
Xantichthys mento 0.12
Family Ostraciidae 0.84
Lactoria diaphanum 0.84
Family Tetraodontidae 0.72 0.72
Lagocephalus lagocephalus 0.72 0.72
Unidentified fishes 0.14 0.23 0.12
Total number of prey 691 427 1665
* Individual prey could not be enumerated.
Table 4
Taxonomic composition in %O of the prey of common dolphinfish from
five areas of the eastern Pacific Ocean (Fig. 1) and from all areas
combined. The occurrence of hard parts (cephalopod mandibles and
fish otoliths) was included. Area codes are N = north, W = west,
E = east, SW = southwest, SE = southeast.
Area
Taxon N W E
Phylum Mollusca 25.0 15.79 15.52
Class Cephalopoda 25.0 15.79 15.52
Order Teuthoidea 25.0 10.53 15.52
Family Enoploteuthidae 2.63
Abraliopsis falco 2.63
Family Mastigoteuthidae 0.86
Mastigoteuthis spp. 0.86
Family Ommastrephidae 25.0 7.89 8.62
Dosidicus gigas 7.89 6.03
Sthenoteuthis oualaniensis 25.0 4.31
Family Onychoteuthidae 9.48
Onychoteuthis banksii 9.48
Onychoteuthis spp.
Family Pholidoteuthidae
Pholidoteuthis boschmani
Family Thysanoteuthidae 0.86
Thysanoteuthis rhombus 0.86
Order Octopoda 7.89 0.86
Family Tremoctopodidae
Tremoctopus violaceus
Family Argonautidae 5.26 0.86
Argonauta spp. 5.26 0.86
Family Bolitaenidae 2.63
Japetella diaphana 2.63
Phylum Arthropoda 25.0 8.62
Class Crustacea 25.0 8.62
Order Decapoda 25.0 7.76
Family Galatheidae 25.0
Pleuroncodes planipes 25.0
Family Penaeidae 4.31
Family Portunidae 3.45
Portunus xantusii 3.45
Phylum Chordata 50.0 36.84 43.10
Class Osteichthyes 50.0 36.84 43.10
Order Clupeiformes 0.86
Family Engraulidae 0.86
Order Stomiiformes 0.86
Family Phosichthyidae 0.86
Vinciguerria lucetia 0.86
Order Myctophiformes 25.0 10.53 6.90
Family Myctophidae 25.0 10.53 6.90
Benthosema panamense 2.63
Myctophum aurolaternatum 6.03
Myctophum nitidulum 0.86
Symbolophorus spp. 7.89 3.45
Order Lampridiformes
Family Trachipteridae
Desmodesma polystictum
Order Beloniformes 25.0 13.16 36.21
Family Belonidae 0.86
Strongylura spp. 0.86
Family Exocoetidae 13.16 25.86
Cheilopogon furcatus
Cheilopogon spilonotopterus
Cheilopogon spp. 5.26 1.72
Exocoetus monocirrhus 12.07
Exocoetus volitans 7.89 6.90
Exocoetus spp. 2.59
Hirundichthys speculiger 1.72
Prognichthys spp. 1.72
Family Hemiramphidae 25.0 2.63 24.14
Oxyporhamphus micropterus 25.0 2.63 24.14
Order Perciformes 50.0 17.24
Family Echeneidae
Rhombochirus osteochir
Family Coryphaenidae 50.0 0.86
Coryphaena equiselis 50.0
Coryphaena hippurus
Family Carangidae 1.72
Naucrates ductor 1.72
Family Gempylidae 7.76
Gempylus serpens 7.76
Family Scombridae 5.17
Acanthocybium solandri 0.86
Auxis spp. 2.59
Euthynnus lineatus 0.86
Thunnus spp. 0.86
Family Nomeidae 6.90
Cubiceps pauciradiatus 6.90
Order Tetraodontiformes 25.0 18.42 3.45
Family Balistidae 25.0 5.26
Xantichthys mento 5.26
Family Ostraciidae 10.53
Lactoria diaphanum 10.53
Family Tetraodontidae 25.0 2.63 3.45
Lagocephalus lagocephalus 25.0 2.63 3.45
Unidentified fishes
Total number of samples 4 38 116
Area
Taxon SW SE All
Phylum Mollusca 14.34 45.22 21.28
Class Cephalopoda 13.97 45.22 21.10
Order Teuthoidea 12.13 43.48 19.45
Family Enoploteuthidae 0.37 0.87 0.55
Abraliopsis falco 0.37 0.87 0.55
Family Mastigoteuthidae 0.74 0.55
Mastigoteuthis spp. 0.74 0.55
Family Ommastrephidae 10.29 43.48 16.88
Dosidicus gigas 7.35 37.39 13.39
Sthenoteuthis oualaniensis 2.57 8.70 4.22
Family Onychoteuthidae 1.84 2.94
Onychoteuthis banksii 1.47 2.75
Onychoteuthis spp. 0.37 0.18
Family Pholidoteuthidae 0.74 0.37
Pholidoteuthis boschmani 0.74 0.37
Family Thysanoteuthidae 0.37 0.37
Thysanoteuthis rhombus 0.37 0.37
Order Octopoda 3.68 5.22 3.67
Family Tremoctopodidae 3.48 0.73
Tremoctopus violaceus 3.48 0.73
Family Argonautidae 3.68 1.74 2.75
Argonauta spp. 3.68 1.74 2.75
Family Bolitaenidae 0.74 0.55
Japetella diaphana 0.74 0.55
Phylum Arthropoda 1.10 2.75
Class Crustacea 1.10 2.75
Order Decapoda 0.37 2.02
Family Galatheidae 0.18
Pleuroncodes planipes 0.18
Family Penaeidae 0.92
Family Portunidae 0.73
Portunus xantusii 0.73
Phylum Chordata 42.28 25.22 38.53
Class Osteichthyes 42.28 25.22 38.53
Order Clupeiformes 0.18
Family Engraulidae 0.18
Order Stomiiformes 2.21 1.28
Family Phosichthyidae 2.21 1.28
Vinciguerria lucetia 2.21 1.28
Order Myctophiformes 0.37 2.57
Family Myctophidae 0.37 2.57
Benthosema panamense 0.18
Myctophum aurolaternatum 1.28
Myctophum nitidulum 0.18
Symbolophorus spp. 0.37 1.47
Order Lampridiformes 0.74 0.37
Family Trachipteridae 0.74 0.37
Desmodesma polystictum 0.74 0.37
Order Beloniformes 32.35 20.00 29.17
Family Belonidae 0.87 0.37
Strongylura spp. 0.87 0.37
Family Exocoetidae 28.68 19.13 24.77
Cheilopogon furcatus 0.74 0.37
Cheilopogon spilonotopterus 0.37 0.18
Cheilopogon spp. 4.04 4.35 3.67
Exocoetus monocirrhus 0.74 0.87 3.12
Exocoetus volitans 18.01 6.09 12.29
Exocoetus spp. 6.99 9.57 6.06
Hirundichthys speculiger 0.87 0.55
Prognichthys spp. 0.74 0.73
Family Hemiramphidae 6.99 8.99
Oxyporhamphus micropterus 6.99 8.99
Order Perciformes 11.40 5.22 10.83
Family Echeneidae 0.37 0.18
Rhombochirus osteochir 0.37 0.18
Family Coryphaenidae 1.84 1.74 1.83
Coryphaena equiselis 0.37 0.55
Coryphaena hippurus 0.74 0.37
Family Carangidae 1.84 1.28
Naucrates ductor 1.47 1.10
Family Gempylidae 1.47 0.87 2.57
Gempylus serpens 1.47 0.87 2.57
Family Scombridae 4.41 2.61 3.85
Acanthocybium solandri 0.18
Auxis spp. 4.04 2.57
Euthynnus lineatus 0.18
Thunnus spp. 0.37 2.61 0.92
Family Nomeidae 1.84 2.39
Cubiceps pauciradiatus 1.84 2.39
Order Tetraodontiformes 1.47 2.94
Family Balistidae 0.55
Xantichthys mento 0.37
Family Ostraciidae 0.73
Lactoria diaphanum 0.73
Family Tetraodontidae 1.47 1.83
Lagocephalus lagocephalus 1.47 1.83
Unidentified fishes 0.74 0.87 1.10
Total number of samples 272 115 545